1944. Ruth Aspinall loves flying and soon becomes an accomplished pilot. When war breaks out and the Air Transport Auxiliary is formed she is anxious to join them. In America, brother and sister, Jack and Lucy Nelson, experienced pilots, make their way to England, also determined to become part of the ATA. Jack and Ruth meet, but with danger all around them, Jack won't commit himself to a serious relationship. By D-Day, Jack has a Spitfire to deliver to France, but his plane never arrives. Ruth is heartbroken but cannot accept that she will never see Jack again.
